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Abstract Temperature and water stress are important factors limiting the gross primary productivity (GPP) in terrestrial ecosystems, yet the extent of their influence across ecosystems remains uncertain. This study examines how surface air temperature, soil water availability (SWA) and vapor pressure deficit (VPD) influence ecosystem light use efficiency (LUE), a critical metric for assessing GPP, across different ecosystems and climatic zones at 80 flux tower sites based on in situ measurements and data assimilation products. Results indicate that LUE increases with temperature in spring, with higher correlation coefficients in colder regions (0.79–0.82) than in warmer regions (0.68–0.78). LUE reaches a plateau earlier in the season in warmer regions. LUE variations in summer are mainly driven by SWA, exhibiting a positive correlation indicative of a water‐limited regime. The relationship between the daily LUE and daytime temperature shows a clear seasonal hysteresis at many sites, with a higher LUE in spring than in fall under the same temperature, likely resulting from younger leaves being more efficient in photosynthesis. Drought stress influences LUE through SWA in all ranges of water availability; VPD variation under moderate conditions does not have a clear influence on LUE, but extremely high VPD (exceeding the threshold of 1.6 kPa, often observed during extreme drought‐heat events) causes a dramatic reduction of LUE. Our findings provide insight into how ecosystem productivities respond to climate variability and how they may change under the influence of more frequent and severe heat and drought events projected for the future. 

Abstract South America, especially the Amazon region, is considered a hotspot of biosphere–atmosphere interactions and presents a unique challenge for regional climate modeling. Here, we evaluate the performance of a regional model in simulating the climate–vegetation system in South America and use the model to investigate the potential role of large‐scale warming in the recently observed trend of hydroclimate and vegetation. Compared with prescribing vegetation based on observational data, adding the predictive vegetation capacity to the regional climate model enabled the model to simulate the vegetation response to climate while sustaining the model performance in reproducing the mean, variability and extremes of the regional climate. The coupled vegetation–climate model captures the recent trends in hydroclimate and vegetation productivity and their spatial contrasts, including a trend toward warmer, drier, and less productive conditions in the Amazon and Nordeste regions and a trend toward cooler, wetter, and more productive condition in the La Plata region. Results from sensitivity experiment driven by detrended boundary forcing for the regional climate suggest that much of the trends in the Amazon and Nordeste regions can be attributed to the effects of large‐scale warming, but contribution from decadal variability also plays a role especially for the most recent decade. However, the trend in the La Plata region cannot be attenuated by the removal of the boundary forcing trend, indicating the role of large‐scale circulation pattern changes. The recent trends in vegetation productivity may be early manifestation of future changes in the Amazon and surrounding regions.